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In Madurai, veteran rooster trainer Periyasamy aka "Pettaikaaran" and Madurai Central Police Inspector Rathnaswamy are tough competitors in cockfights, and it is often Pettai who wins because nobody knows his way of maintaining the quality of the birds. Rathnasamy and Pettai were once protegees of grand veteran Doraisamy, who has retired from active competition but is considered as the honorary overlord of rooster fighting in South Tamil Nadu. K. P. Karuppu and Durai are the favourites in Pettai's team. Karuppu is very talented in breeding and training roosters, while Durai is very rich, owns three bars in the town of Thiruparankundram, and is talented in rooster training. One day, during a informal match in rural field area, local police raid the spot and arrest most of the involved, except Rathnaswamy and Durai who escapes. Everyone is released after stern warning from the local police, after intervention of Rathnasamy. Pettaikaran's roosters were abducted by the fleeing men from Rathnasamy's party, and Pettaikaran orders his guys to bring them back by any means. He also asks them to kill any fowl which has bred with his roosters, to prevent the strength of his roosters passing to Rathnasamy's flock. Karuppu goes to the railway colony at night, to retrieve their rooster, since an ally of Rathnasamy lives there. He kills the fowl which had mated with their rooster, and bumps into Anglo-Indian girl named Irene. He is smitted by her beauty, but silences her and leaves. Having been defeated in all earlier bouts with Pettaikaran's roosters, Rathinasamy keeps insisting on having one last fight to win and satisfy his old mother's wish, but Pettai declines to have any more fights with him because he feels that Rathnaswamy has lost faith in his roosters and will be using nefarious methods to win, like the earlier raid. Karuppu falls in love with Irene, who dislikes him. He does not recognize this initially and keeps following her. One day, the residents of Irene's colony confront him and ask Irene to tell who she is in love with – Karuppu or Dinesh, another man in the locality. She points her finger at Karuppu, and he goes into rapture. She explains later that she had to lie in order to avoid Dinesh, who has been bothering her for a long time. Later, Irene too falls in love with Karuppu. Meanwhile, Rathnasamy tries to cajole Ayub, Pettai's veterinarian for roosters, to get Pettai to come for a competition against himself, in exchange for a heavy amount. Ayub refuses and insults Rathnasamy. The same night, Ayub is killed in a hit and run accident. Durai initially suspects Rathnaswamy and all of Pettai's gang wants revenge, but Pettai decides to conduct a state tournament in Ayub's name and provide his family with funds so that Ayub's poor daughters can be married. He asks permission from Rathnasamy as the police have to permit it, but Rathnaswamy refuses, manhandles, and berates Pettai as a thief. In a rage, Pettai, suspecting Rathnaswamy of the murder, agrees to the one-on-one rooster fight that Rathnaswamy has been asking for so long. He bets that he will field his roosters against every rooster that Rathnaswamy brings to the field in the following tournament, and even if one of Pettai's roosters loses against Rathnaswamy's, Pettai will tonsure his head and face, publicly apologize to him, and will quit rooster fights. If Rathnaswamy is not able to beat at least one of his roosters, the same conditions will be applied on him, at the end of the day. Finally getting his way, Rathnaswamy permits the tournament. The grand state tournament is arranged by Pettai's team, getting heavy funds and official permission. In the initial 11 fights, Rathnasamy's roosters are defeated by Pettai's. Rathnasamy brings high-bred roosters from Bangalore and enters them into the tournament. Seeing the quality of the new roosters, Karuppu asks Pettai to let his rooster fight, but Pettai puts his rooster down and says that he will choose the best-competing rooster himself. Karuppu surreptitiously enters the contest as Pettai's team before Pettai can choose the rooster because he needs to repay Irene for the loan he took to prepare his rooster for the fight. Pettai does not believe in Karuppu and his rooster announces that Karuppu will not represent Pettai, and the result of Karuppu's match will not be acknowledged by his team. However, the 'underdog' emerges victorious in three consecutive rounds, despite facing roosters spiked with steroids. Karuppu gets the best coach award of the tournament and his bet money of ₹10 lakh (equivalent to ₹21 lakh or US$25,000 in 2023).


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